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Division of Integrative Organismal Systems

IOS Collaborators and Other Affiliations (COA) Template

The Collaborators and Other Affiliations (COA) template, available for download here as an Excel file, is required for all IOS preliminary and full proposals. 

For information about completing and submitting the COA template, as well as troubleshooting advice, please read this post on the IOS blog.

Last updated: Dec 20, 2016

FAQs for IOS Core Programs solicitation

Frequently Asked Questions for revised IOS Core Programs solicitation (NSF 17-508):

See also the IOS blog:

Name Email Phone Room
Behavioral Systems  Cluster (703) 292-8423  685N  
Developmental Systems  Cluster (703) 292-8417  685N  
Neural Systems  Cluster (703) 292-8421  685N  
Phys. & Struct. Systems  Cluster (703) 292-8413  685N  


Solicitation  17-508

Important Information for Proposers

A revised version of the NSF Proposal & Award Policies & Procedures Guide (PAPPG) (NSF 17-1), is effective for proposals submitted, or due, on or after January 30, 2017. Please be advised that, depending on the specified due date, the guidelines contained in NSF 17-1 may apply to proposals submitted in response to this funding opportunity.


Full Proposal Deadline Date

    August 4, 2017

    First Friday in August, Annually Thereafter

Preliminary Proposal Deadline Date

    January 19, 2018

    Third Friday in January, Annually Thereafter




The Division of Integrative Organismal Systems (IOS) supports research aimed at understanding why organisms are structured the way they are and function as they do. Proposals should focus on organisms as a fundamental unit of biological organization. Principal Investigators (PIs) are encouraged to apply systems approaches that will lead to conceptual and theoretical insights and predictions about emergent organismal properties. Areas of inquiry include, but are not limited to, developmental biology and the evolution of developmental processes, nervous system development, structure, and function, physiological processes, functional morphology, symbioses, interactions of organisms with biotic and abiotic environments, and animal behavior.

Proposals are welcomed in all of the core scientific program areas supported by the Division of Integrative Organismal Systems.  All investigator-initiated proposals submitted to this solicitation must be invited based on merit review of preliminary proposals. There is a single submission deadline with a limit of 2 preliminary proposals per investigator per year as PI or Co-PI in response to this solicitation. Please see the PAPPG for definition of roles for PI and Co-PI. The PI/Co-PI limits apply only to the preliminary proposals submitted to this solicitation and do not pertain to proposals submitted in response to other IOS or NSF solicitations.


Behavioral Systems

Developmental Systems

Neural Systems

Physiological and Structural Systems

Division of Integrative Organismal Systems Core Programs

What Has Been Funded (Recent Awards Made Through This Program, with Abstracts)

Map of Recent Awards Made Through This Program